BULLETINS OF AMERICAN PALEONTOLOGY
STUDIES IN PALEONTOLOGY
Professor of Geology, University of Cincinnati
ON HIS 4Sth year of teaching.
BULLETINS OF AMERICAN PALEONTOLOGY
No. 287, 1975
STUDIES IN PALEONTOLOGY
U.S. Geological Survey
Washington, D.C. 20244
John K. Pope
Oxford, Ohio 45056
affectionately dedicated to
Library of Congress Card Number: 75-1
Printed in the United States of America
Arnold Printing Corporation
Biography of K. E. Caster
La Casa en
publications of K. E. Caster
and paleoecology of Carboniferous
Group, Brooks Range, Arctic Alaska
Augustus K. Armstrong
Ontogeny and systematics of Timeischytes
matic Devonian edrioasteroid
Bruce M. Bell
Podial efficacy of some Ordovician asteroids (Echinodermata) from
Lower Carboniferous brachiopods from Axel Heiberg and Melville
Islands, Canadian Arctic Archipelago
Middle Devonian Bone Beds and the Columbus-Delaware (Onondagan-Hamiltonian) contact in Central Ohio
James E. Conkin and Barbara M. Conkin
bifoliate tubular bryozoan genera from the Simpson Group
(Middle Ordovician), Arbuckle Mountains, Oklahoma
George T. Farmer, Jr
American Lituitidae (Cephalopoda)
Rousseau H. Flower
Some problems in coral phylogeny and classification
Rousseau H. Flower and Helen M. Duncan
Maquoketa-like molluscan community in the Brassfield Formation
(Early Silurian) of Adams County, Ohio
William B. Harrison, III and Linda Kelley Harrison
new late Cretaceous generic link in limulid lineage
Mark Erickson and Douglas E. O'Brien
F. D. Holland, Jr., J.
Ontogenies of three late Cambrian trilobites from the
Formation, northern Black Hills, South Dakota
ancestry, geographical extent, and fate of the Brassfield coral
fauna (Middle Llandovery, North America)
Richard S. Laub
Hitchhiking clams in the Marcellus Sea
Osborne B. Nye, Jr., James C. Brovver and Steven E. Wilson
Trilobite trace fossils from the Clinton
Central New York State
Richard G. Osgood,
and William T. Drennen, HI
Systematics and functional morphology of Columbocystis, a Middle
Ordovician "Cystidean" (Echinodermata) of uncertain affinities
Ronald L. Parsley
Fordilla iroyensis Barrande and early pelecypod phylogeny
John Pojeta, Jr
Evidence for relating the Lepidocoleidae, machaeridian echinoderms,
to the mitrate carpoids
John K. Pope
The genus Plumalina
Hall, 1858 (Coelenterata)
Daniel B. Sass and Barrett N. Rock
Monocyclism vs. Dicyclism:
John M. Warn
sexual dimorphism and ontogeny of Ceratopsis chambersi (Mil(Ostracoda, Palaeocopida) from the Upper Ordovician of southwestern Ohio and northern Kentucky
Steven M. Warshauer
STUDIES IN PALEONTOLOGY
KENNETH EDWARD CASTER
an art as well as a science."
Kenneth Caster is a complex man. Raised and educated in upNew York, and Pennsylvania, he blends the resourcefulness
and craftsmanship of the Yankee pioneers of his part of this country
with the imagination and analytical mind of the scientist and the
wit and sophistication of a man of letters and international travel.
Those who have learned from him have come away inspired, aware
not only of their science but also of the breadth of man's other finer
endeavors, such as art and literature, and of nature's riches. Few scientists combine the penetrating intelligence, rapid comprehension of
detail, stamina, and enthusiasm for life to permit themselves simultaneously to become both masters of their chosen fields and broadly
authoritative throughout the arts, humanities, and other sciences as
Born in New Albany, Pennsylvania, in 1908, Kenneth Caster
grew up in Ithaca, New York. There he received his early education and at Cornell University, his A.B. in 1929, M.S. in 1931, and
Ph.D. in 1933. Even before entering college, Kenneth came under
the influence and inspiration of J. C. Bradley, entomologist, and
E. Laurence Palmer, natural historian, both of Cornell University.
In his early college years. Caster's interest was entomology.
even while serving as an assistant
to Bradley, his interest was diverted toward the work of G. D.
Harris, paleontologist and stratigrapher in the Department of
ever, as a senior,
Geology at Cornell University. Harris, although a
Tertiary mollusks, recognized the need for continuing work in the
New York and there
The result was Caster's study of the
and stratigraphy of the Upper Devonian and Lower Carboniof southwestern New York and northwestern Pennsylvania,
mid-Paleozoic rocks of Pennsylvania and
directed Caster's attention.
his doctoral dissertation, his early publications,
interest in that part of the geologic
a graduate student at
Caster served as
in the post of Instructor of Paleontology.
receiving his doctoral degree, and after his marriage to the
gracious Anneliese, Kenneth
rapidly through the positions of
Instructor of Geology and Paleontology at Cornell, and Assistant
of Science at the
arrived at the University of Cincinnati in 1936 as Instructor in
Geology and Curator
Geology Museum and rose steadily to
Geology (1952) and Fellow of
his present position of Professor of
the Graduate School. Additionally, he held visiting professorships at
Potsdam State College (1936), Universidade de Sao Paulo, Brazil
(1944-47), Escuela Nacional de Minas, Colombia (1948), University
of Virginia Biological Station (1955), University of Tasmania (195657), University of California at Berkeley (1962), University of
Germany (1964), and
University of Southern California
Kenneth's society memberships,
distinctions are too
and even honors and
recipient of the
Derby Medal at the Brazihan Geological Survey Centennial (1952)
and of the Gondwana Medal at the Geological Survey of India
Centennial (1956). He has received Guggenheim and Fulbright Fellowships for travel and study in South America, Africa, Australia,
Zealand and grants from the National Research Council,
Geological Society of America and National Science Foundation for
study of Upper Ordovician, Devonian, and Mississippian faunas and
stratigraphy. He is a fellow or has been an executive officer of ten
of the 38 societies
is a member, inMacDowell Society (1964-
cluding the presidency of the Cincinnati
66), Paleontological Society (1960), and twice the presidency of the
Paleontological Research Institution.
find the source of Caster's success
the impressively long
of geological time
and acclaim, we must scan
Their subjects span
of his publications.
and touch upon almost
glance, there seems to be no general systematic or chrono-
logic organization to the papers. It quickly
becomes apparent that
the papers are divided into vaguely defined periods. Like the paintings of Picasso, Caster's papers are arranged into periods reflecting
changing interests: the Devono-Mississippian Period, by exten-
Hemisphere Period, the Sponge Period, the Jellyfish Period, the Merostome Period, the Continental Drift Period and
the Echinoderm-Carpoid Period, The papers may also be organized
by theme. Some, as the Devono-Mississippian faunal and stratigraphic studies and the South American studies, are comprehensive
attacks upon large and widely known geological problems. Others,
as the merostome and carpoid papers, began with serendipity, the
sion, the Austral
chance discovery of
out of place in time or of such exceptional
preservation as to shed
of papers in this category are Caster building
by Caster. Underlying
of the papers
excellence in comprehension and presentation,
imagination in finding and exposing the unique qualities of each
Moreover Caster dared
to be unconventional.
stration of facies in his early work, at that time a novel
unpopular concept, has become a
His eloquent insistence over
years that continental drift
"a topic at least worthy of consideration" has finally been justified.
however, do not reveal the
a trustee of the
he and Anneliese regularly
have attended musical and theater productions
Kenneth's other interests that his personality
Playhouse in the Park. Kenneth delights
handsome home displays not only the
and handicrafts which are mementos of their global
but also endless bookshelves housing their non-scientific collection.
Kenneth's spacious office and anteroom at the University contain
one of the finest private paleontological libraries in the world. His
voluminous and international. From
secondary benefit, he adds to his extensive stamp collection. Caster's
tolerance of students'
legendary, as witnessed
from the "University of the State of Collapse", his
charter membership in the "Lipalian Research Foundation", his
"Agawana Medal", and his perseverance under the stare from the
bronze bust of Ernst Haeckel placed high atop a bookcase in his
generous with his time and information. Years
ago, there clustered about
of dedicated, enthusiastic,
organized to become the
In return for patient instruction, he has received from them some
of the finer specimens in the
Geology Museum. Both Kenneth and
AnneUese are warm, generous, and hospitable.
ings of graduate students at their
their continuing concern
for students has resulted in a binding, life-long esprit
de corps. Not
only Kenneth's mastery of paleontology, his outstanding teaching
and research, but also his magnetic personality have drawn generations of students to Cincinnati.
LA CASA EN CLIFTON HEIGHTS
El homenaje a un cientifico que llega a los 45 anos de vida, no
solo de sobresalientes trabajos de investigacion, sino de
universitaria para cuyo desempeiio feliz se necesita
monacal, no puede ser mas merecido y oportuno.
Hay sinembargo, fuera de la parte cientlfica,
que aglutina en un todo coherente,
hombre. Quien esto escribe ha tenido
amistad de Kenneth y
Ana Caster por mas
desea rememorar en estas cortas lineas, la trayectoria del
Caster como un lugar siempre abierto para
y extranjeros conocidos a
largo de los extensos viajes de los Caster por Sur America, Africa,
Australia, Europa, etc.
— Aquella casona de Cincinnati ha llegado a
Embajada diplomatica sin el pesado aspecto
con una incomparable y acogedora llaneza que
hospitalidad de los tiempos coloniales norteamericanos.
Como uno de los paleontologos extranjeros que se ha acogido
muchas veces al techo protector y hospitalaria de Kenneth y Ana,
me atrevo a hacerme vocero de los muchos otros visitantes cientificos
multinacionales, al pedir a los espiritus protectores de la Paleontologia
sus cultores, permitan
muchos anos por
hogar de Clifton Heights y sus duenos, sigan siendo el lugar obligado de peregrinacion bajo la sombra del arioso Gingko, arbol
mas que apropiado para proteger
doble caracter de arbol sagrado
hogar de un paleontologo, en su
THE PUBLICATIONS OF KENNETH
faunas of the Upper Allegheny. Bull. Amer. Paleont, vol,
174 pp., 59 pis.
Upper Devonian rocks at Ithaca, Neio York. (Abstract), Geol. Soc.
Bull., vol. 44, pp. 201-202.
Stratigraphic relationships in northtwestern Pennsylvania. (Abstract).
Geol. Soc. America, Bull., vol. 44, pp. 202-203.
Fades nomenclature and the Upper De-vonian. (Abstract). Geol. Soc.
America, Proc. 1933, pp. 348-349.
The stratigraphy and paleontology of northiuestern Pennsylvania, Part 1,
Stratigraphy. Bull. Amer. Paleont., vol. 21, No. 71, 185 pp., 2 pis.
With P. D. Torrey and G. H. Chadwick. Demise of the Bradfordian
Series. Amer. Assoc. Petroleum Geol., Bull., vol. 19, pp. 912-915.
Upper Devonian marine faunas in the Seaivard Phase of the Catskill
magnafacies. (Abstract). Geol. Soc, America, Proc. 1934, p. 363.
Boundary between the Devonian and Mississippian Systems in ivestern
Pennsylvania. (Abstract). Geol. Soc. America, Proc. 1934, pp. 441-442.
With A. A. Olsson. Occurrence of Baculiies ovatus Zone of Upper Alberta
Shales in southeastern British Columbia. Amer. Assoc. Petroleum Geo!.,
Bull., vol. 19, pp. 295-299; (Abstract). Geol. Soc. America, Proc. 1934, pp,
With Bradford Willard. Age of the Devonian of south^vestem Pennsylvania. Amer. Assoc. Petroleum Geol., Bull., vol. 19, pp. 1546-1550.
With R. H. Flower. The stratigraphy and paleontology of northiuestern
Pennsylvania, Part 2, Paleontology, Section A, The Cephalopod Fauna
of the Coneivango Series of the Upper Devonian in Nev: York and
Pennsylvania. Bull. Amer. Paleont, vol. 22, No. 75, pp. 199-270, 8 pis,
With A. A. Olsson. Devonian fauna from Colombia, South America.
(Abstract). Geol. Soc. America, Proc. 1936, pp. 369-370.
A restudy of the tracks of Paramphibius. Jour. Paleont., vol. 12, pp. 3-60,
13 pis. (Abstract). Geol. Soc. America, Proc. 1936, pp. 366, 1937.
Nezu genera of Upper Devonian Pelccypoda. (Abstract). Geol. Soc.
America, Proc. 1937, p. 271, 1938.
Some facieological aspects of the pelecypod fauna of the Coneivango
Series of the Upper Devonian in New York and Pennsylvania. (Abstract).
Geol. Soc. America, Proc. 1937, pp. 271-272, 1938.
Pterygotus in the Richmond of Ohio. (Abstract). Geol. Soc. America,
vol. 49, p.
Preliminary notes on the revision of the crenulate-hinged Strophomenacea.
(Abstract). Geol. Soc. America, Bull., vol. 49, p. 1911.
1938. Macroscopic fauna of the Ouimbriz (Eocene) Formation on the Luculo
River, Angola. Communicacoes dos Services Geologicos de Portugal, vol.
20, pp. 1-37.
Siliceous sponges from the Mississippian and Devonian strata of the
Penn-York Embayment. Jour. Paleont., vol. 13, pp. 1-20, 4 pis. (Abstract).
Geol. Soc. America, Bull., vol 49, pp. 1910-1911, 1938.
Comparison of the siliceous sponges Armstrongia Clarke, 1920, and
Titusvillia Caster, 1939. Jour. Paleont., vol. 13, pp. 531-532.
Were Micrichnus scotti Abel and Artiodactylus sinclairi Abel of the
Newark Series (Triassic) made by vertebrates or limuloidsf Am. Jour.
Sci., vol. 237, pp. 786-797.
Devonian fauna from Colombia.
Paleont., vol. 24, No. 83,
pp., 14 pis.
vertebrates make the trails Kouphichnium (Jurassic) and Micrichnus
(Triassic)? (Abstract). Geol. Soc. America, Bull., vol. 50, p. 1971.
Penn-York Embayment. (Abstract). Geol.
stratigraphy of the
Bull., vol. 50, pp.
With W. H. Bucher and Stewart Jones. Elementary description
cinnatian fossils and strata and plates of commoner fossils in the
Die sogenannten "Wirbeltierspuren" und die Limulusfdhrten der Soln-
of Cincinnati, Ohio. Univ. Cincinnati, 13 pp., 8 pis.
hofencr Plattenkalkc. Palaeontologische Zeitschrift, vol. 22, pp. 12-29.
The Titusi/illiidae; Paleozoic and Recent branching Hexactinellida.
Palaeontographica Americana, vol. 2, No. 12, 52 pp., 5 pis.
1941. Trails of Limulus and supposed vertebrates from Solnhofen Lithographic
Limestone. Pan-American Geologist, vol. 76, pp. 241-258.
1941. Paleozoic geography of South America ivith particular reference to the
Devonian. (Abstract). Geol. Soc. America, Bull., vol. 52, p. 1892.
1941. Titusvilliidae: Useful stratigraphic indices. (Abstract). Geol. Soc. Amer1941.
ica, Bull., vol. 52, p. 1962.
from the Upper Cambrian of IVyoming. Amer. Jour. Sci.,
pp. 104-112, pi. 1; (Abstract). Geol. Soc. America, Bull., vol.
52, pp. 1962, 1941.
The paleontological collections of the University of Cincinnati. Science,
No. 2470, pp. 454-455.
With G. A. Cooper,
Formations of North America. Geol. Soc. America,
Bull., vol. 53, pp. 1729-
1794, 1 pi.
The age and relations of the Colombian Devonian strata. Eighth American
Science Congress (Washington, D.C., 1940), Proc, vol. 4, pp. 27-67.
Tvi)o siphonophores from the Paleozoic. Palaeontographica Americana,
vol. 3, No. 14, 34 pp., 2 pis.
Limulid trails from the Upper Triassic (Chinle) of the Petrified Forest
National Monument, Arizona. Amer. Jour. Sci., vol. 242, pp. 74-84, pi. 1.
Hydrozoan jellyfish from the Loiver Cretaceous of Texas. (Abstract).
Geol. Soc. America, Bull., vol. 55, p. 1465.
New synziphosuroid merostome from the Loiver Ordovician of Tennessee.
(Abstract). Geol. Soc. America, Bull., vol. 55, pp. 1465-1466.
fVas Rhizostomites admirandus Haeckel of Solnhofen a true jellyfish?
(Abstract). Geol. Soc. America, Bull., vol. 55, p. 1466.
A new jellyfish (Kirklandia texana Caster) from the Loiver Cretaceous
of Texas. Palaeontographica Americana, vol. 3, No. 18, 52 pp., 5 pis.
Ncvu names for tivo homonyms. Jour. Paleont., vol. 19, p. 319.
With W. H. Bucher and S. M. Jones. Elementary guide to the fossils and
strata in the vicinity of Cincinnati. Cincinnati Museum Nat. Hist, 31
pp., 9 pis.
Carboniferous deposits of southern Goias and Mato Grosso. (Abstract).
Geol. Soc. America, Bull., vol. 58, pp. 1171-1172.
1947. Devonian System in Goias and Mato Grosso, Brazil. (Abstract). Geol.
Soc. America, Bull., vol. 58, p. 1172.
1947. With S. Petril [Petri]. Devonian stratigraphy and paleontology of the
States of Parana and Sao Paulo, Brazil. (Abstract). Geol. Soc. America,
Bull., vol. 58, p. 1173.
Expedi^do geologica em Goias
Grosso. Mineragao e Metalurgia,
vol. 12, pp. 126-127.
Excursdo geologica ao Estado do Paiu'i (Brasil). Ibid, pp. 271-272.
With J. C. Mendes. DuToifs geological comparison of South America
•with South Africa after tiventy years. (Abstract). Geol. Soc. America,
et al. Correlation of the Misstssippian Formations
of North America. Geol. Soc. America, Bull., vol. 59, pp. 91-196.
Concerning the eurypterid Megaloanachronism. (Abstract). Geol. Soc.
Bull., vol. 62, pp. 1428-1429.
With O. D. von Engeln. Geology, 730 pp.. New York, McGraw-Hill
Stratigraphic and paleontologic data relevant to the problem of AfroAmerican ligation during the Paleozoic and early Mcsozoic. Amer. Mus.
Nat. Hist., Bull., vol. 99, pp. 105-158.
Concerning Enoploura of the Upper Ordovician and its relation to other
carpoid Echinodermata. Bull. Amer. Paleont., vol. 34, No. 141, 56 pp.,
1952. With W. B. Macke. An aglaspid merostome from the Upper Ordovician
of Ohio. Jour. Paleont., vol. 26, pp. 753-757, pi. 109.
1952. Relationships of the carpoid echinoderms in the light of Enoploura. (Abstract). Geol. Soc. America, Bull., vol. 63, p. 1239.
1953. Translator with J. C. Mendes. Comparacao Geologica entre a America
do Sul e a Africa do Sul, by A. L. DuToit. Rio de Janeiro, Inst. Brasil
Geog. e Estatistica, Serv. Graf., 179 pp.
Afro-American linkage in the Paleozoic.
Cong., 19th, Algiers, 1952, Comptes Rendus,
sec. 2, fasc. 2, p. 103.
E. N. Kjellesvig-Waering. Melbournopterus, a netv Silurian eurypfrom Australia. Jour. Paleont., vol. 27, pp. 153-156, pi. 20.
1954. Brazilian Devonian carpoids. (Abstract). Jour. Paleont., vol. 28, p. 511.
1954. A new carpoid echinodcrm from the Parana Devonian. Acad. Brasil
de Cien, Anais, vol. 26, pp. 123-147.
Introductory survey of the Brazilian Carboniferous. Bull. Amer. Paleont.,
No. 149, pp. 1-14.
E. A. Dalve and J. K. Pope. Elementary guide to the fossils and
strata of the Ordovician in the vicinity of Cincinnati, Ohio. Cincinnati
Mus. Nat. Hist., 47 pp., 8 pis. New Edition 1961.
Devonian placocystoid echinoderm from Parana, Brazil. In Paleontologia do Parana, F. W. Lange, ed., Curitiba, Brazil, pp. 137-148, pi. 8.
With E. N. Kjellesvig-Waering. Marsupipterus, an unusual eurypterid
from the Doivntonian of England. Jour. Paleont., vol. 29, pp. 1040-1041.
With E. N. Kjellesvig-Waering. The Pterygotidae of the Silurian Vernon
Shales of Neiv York. Jour. Paleont., vol. 29, pp. 1041-1047.
With E. N. Kjellesvig-Waering. Some notes on the genus Dolichopterus
Hall. Jour. Paleont., vol. 30, pp. 19-28, pi. 4.
K. Brooks. Nevj fossils from the Canadian-Chazyan (Ordovician) hiatus in Tennessee. Bull. Am. Paleont., vol. 36, No. 157, pp. 157199, pis. 12-21.
With H. K. Brooks. Pseudoarctolepis sharpei, n. gen., n. sp. (Phyllocarida), from the JVheeler Shale (Middle Cambrian) of Utah. Jour.
Paleont., vol. 30, pp. 9-14, pi. 2.
With T. H. Eaton, Jr. Microstructure of the Plates in the carpoid echinoderm Paranacystis. Jour. Paleont., vol. 30, pp. 611-614, pi. 74.
Australian carpoid echinoderms. (Abstract). Resumes de los Trabajos
International Geological Congress, Mexico, p. 116.
With E. D. Gill and P. C. R. Boswell. Simple apparatus for vacuum injection of moulding latex. Australian Jour. Science, vol. 18^ pp. 198-199.
Annotated Bibliography. In Ladd, H. S., ed., PaleoProblcmatica
ecology. Geol. Soc. America, Mem. 67, pp. 1025-1032.
With E. D. Gill. Australian and New Zealand Silurian-Devonian carpoids. (Abstract). Geol. Soc. America, Bull., vol. 68, p. 1707.
Palaeogeography. Encyclopaedia Americana, vol. 21, pp. 159-163.
Gondwanaland. Encyclopaedia Britannica, vol. 10, pp. 569-570.
With E. D. Gill. Carpoid echinoderms from the Silurian and Devonian of
Australia. Bull. Amer. Paleont., vol. 41, pp. 1-71, 10 pis.
1960. With J. K. Pope. Morphology and affinities of Eocrinus, an archetype
of the Echinodermata. (Abstract). Geol. Soc. America, Bull., vol. 71, pp.
1961. Paleontologica enigtnatica Caster. Jour. Paleont., vol. 35, pp. 238-240.
1961. Sunday all-day excursion in the Cincinnati Region. Field Trip 7, in Geol.
Soc. America Guidebook for Field Trips, Cincinnati Meeting: New York,
Geol. Soc. America, pp. 253-257.
1963. With A. J. Boucot, David Ives, and J. A. Talent. Relationships of a
neiu Loiucr Devonian tcrcbratuloid (Brachiopoda) from Antarctica. Bull.
Amer. Paleont., vol. 46, No. 207, pp. 80-151, pis. 16-41.
neiu class of Echinoderms. Sci1963. With J. W. Durham. Helicoplacoidca:
ence, vol. 140, No. 3568, pp. 820-822.
1964. With E. N. Kjellesvig-Waering. Upper Ordovician curypterids of Ohio.
Palaeontographica Americana, vol. 4, No. 32, pp. 301-358, pis. 43-53.
1965. With R. L. Parsley. North American Soluta (Carpoidea, Echinodermata).
Bull. Amer. Paleont., vol. 49, No. 221, pp. 109-174, pis. 16-18.
1965. Memorial to Ray S. Basslcr (1878-1961). Geol. Soc. America, Bull., vol.
76, pp. 167-174.
1966. With J. W. Durham. Helicoplacoids. In Treatise on Invertebrate
Geol. Soc. America and
Pt. U, Echinodermata 3, v. 1
Univ. Kansas Press, pp. U131-U136.
Pt. S, Echino1967. Homoiostelea. In Treatise on Invertebrate Paleontology
dermata 1, V. 2: Geol. Soc. America and Univ. Kansas Press, pp. S581S627.
idea of a Caster Festschrift occurred to us in the
Kenneth Caster's scholarship in paleonand
the fact that he has had over 50 gradtology and stratigraphy,
uate students more than half of whom obtained their Ph.D. degrees
1972, as a
felt that an anniversary publicawould be his 45th year of teaching.
his charming wife Annie, and Katherine
VanWinkle Palmer, his longtime friend and editor of the publications
of the Paleontological Research Institution, and received enthusiastic
at the University of Cincinnati.
would be appropriate
We first approached
support for the project.
to publish this
book we decided to approach
K.E.C.'s present and former students, friends, paleontological and
Geology at the University
of Cincinnati, and the Dry Dredgers, an organization of collectors
of fossils in and around Cincinnati, Ohio. The response to our request was overwhelming and in short order we had funds enough to
go ahead with the Caster volume. A list of all monetary contributors
faculty colleagues, the
follows this introduction.
cooperation of Katherine Palmer,
the book as a volume of the Bulletins of American
Paleontology, This seemed especially appropriate because
D. Harris, who founded the Bulletins and the
Paleontological Research Institution, and because K. E. C. and his
a student of G.
students have published extensively in the two series of the Paleontological
of paleontology at Cincinnati
the contributed papers assembled here.
subjects include, bio-
morphology, ontogeny, paleoecology, phylogeny, and systematics.
The animal groups discussed are: anthozoans, asteroids, brachiopods,
bryozoans, cephalopods, crinoids, edrioasteroids, gastropods, limulids,
machaeridians, ostracodes, pelecypods, trilobites, and ichnofossils.
Stratigraphically, the papers span the geologic
Cambrian to the Cretaceous, with
column from the
special concentration in the lower
and middle Paleozoic.
K. E. C. has worked with famous colleagues at Cincinnati and
elsewhere and has written
we like to think
number of students who have
from him at Old Tech (the building
for the large
him and learned
which houses the Department of Geology at the University of Cincinnati). Ken Caster's personality, his thorough understanding of
great ability as a teacher, the University's large collec-
tion of fossils,
fact that Cincinnatian rocks
contribute to attracting fledgling students of paleontology
there, students first get a solid
systematics before passing on to other aspects of paleontology.
it was decided to limit the scientific conbook to Ph.D. students of K. E. C; 19 of his students were able to contribute papers. Of those by several writers
only the senior author was a student of Caster. So, Ken, for better
or worse, you have only yourself to blame for what follows, for none
of us would have gotten this far without your help and personal
Early in the project
tributions in this
MONETARY CONTRIBUTORS TO THIS BOOK
A. K. Armstrong
M. G. Arnold (In memory
H. M. Duncan)
G. B. Barbour
Mr. and Mrs. R. L. Black,
College of Wooster
G. Arthur Cooper
East Lansing, Michigan
A. T. Cross
R. A. Davis
H. H. Dresser
at the University
R. H. Durrell
G. T. Farmer
Graduate Fellows University of
C. A. Fleming
R. H. Flower
Bowling Green, Ohio
University of Cincinnati
E. D. Gill
Grand Forks, North Dakota
O. L. Karklins
H. S. Ladd
Grand Forks, North Dakota
Los Angeles, California
Las Vegas, Nevada
Margaret and Martha Neu
New Mexico Institute of
Mining and Technology
O. B. Nye, Jr.
R. G. Osgood
R. L. Parsley
R. A. Pohowsky
John and Mary Lou Pojeta
John K. Pope
P. E. Potter
D. B. Sass
Highland Heights, Kentucky
Irene K. Stephany
Steven and Yvette Warshauer
Morgantown, West Virginia
E. L. Yochelson
STRATIGRAPHY AND PALEOECOLOGY OF
CARBONIFEROUS CORALS, LISBURNE GROUP,
BROOKS RANGE, ARCTIC ALASKA
Augustus K. Armstrong
Menlo Park, California 94025
U.S. Geological Survey,
The Carboniferous Lisburne Group of arctic Alaska contains coral faunas
ranging in age from Osagean (Early Mississippian) to Atokan (Middle
Pennsylvanian). Osagean beds have a small fauna of solitary and tabulate
corals. Meramecian and lowermost Chesterian beds contain a large fauna of
Ekvasophyllum spp., Fabcropliyllum spp., Diphyphyllum 'venosum Armstrong,
D. klaivockensis Armstrong, D. nasorakcnsis Armstrong, Lithostrotion (Siphonodendron) dutroi Armstrong, L. (S.) sinuosum (Kelly), L. (S.) ivarreni Nelson,
L. (S.) lisburnensis Armstrong, Lithostrotion reiscri Armstrong, Lithostrotionella
niakensis Armstrong, L. banffcnsis (Warren), L. mclareni (Sutherland), L.
birdi Armstrong, L. pennsylvanica
(Shimer), Thysanophyllum astraeiforme
(Warren), T. orientale Thomson, Sciop/iyllum lambarti Harker and McLaren,
and S. alaskacnsis Armstrong. Corals, rare in younger Chesterian beds, are
Lithostrotionella aff. L. mclareni (Sutherland), Lithostrotion (S-) ignekensts
Armstrong, Syringopora spp., and occasional solitary corals. Pennsylvanian
(Atokan) beds of the Lisburne Group contain Lithostrotionella ivahooensis Armstrong, Corivenia jagocnsis Armstrong, a thick-walled syringoporoid, and
Preliminary studies of the lithostrotionoids indicate that the Lisburne
Group can be divided into eight coral-assemblage zones. The zones in ascending
order are: Meramecian, 4 zones: Lithostrotion (S.) dutroi Armstrong, L. reiseri
Armstrong, Lithostrotionella mclareni (Sutherland), and Sciophyllum lambarti
Harker and McLaren; Meramecian-Chesterian transition, 1 zone: Lithostrotion
(S.) lisburnensis Armstrong; Chesterian, 2 zones: Lithostrotionella
mclareni (Sutherland) and Lithostrotion (S-) ignekensis Armstrong; Atokan,
1 zone: Corivenia jagoensis Armstrong.
Lisburne limestones were deposited on a slowly subsiding carbonate platform and are cyclic. Colonial corals of Meramecian and Atokan age occur
in carbonates associated with shallow-water shoaling facies. The paucity of
corals in carbonates of Osagean, Chesterian, and Morrowan age is attributed
to regional temperature or salinity changes that inhibited their growth. Carboniferous corals are not known to have formed reefiike masses in arctic Alaska,
but some biostromal accumulations occur.
locations of Carboniferous outcrops in arctic Alaska are
in Text-figure 1. Field collecting
tory studies are only partly completed; therefore, this paper
interim report. Lisburne solitary rugose and tabulate coral faunas
have not been studied and
which does not
poorly represented in this
Thirty-six stratigraphic sections in the Brooks Range, arctic
Alaska, were measured by Jacob's staff and tape.
Rock and foramini3 m). Coral
Thin sections were
samples were collected every 5 to 10 feet (1.5 to
were made throughout the section.
Carboniferous corals, Alaska
carbonate microfacies and microfossil studies.
were studied by means of 2-inch by 3-inch oriented thin sections.
Bernard L. Mamet's microfossil identifications and zonations
are used in the biostratigraphic analysis of the Lisburne
and the determination of the vertical ranges of
wish to express
appreciation to Irvin L. Tailleur, the
and Hillard N. Reiser, the party
summers of 1969-1971, for their generosity in supporting my Lisburne Group coral collecting and stratigraphic studies. I wish to
thank the Naval Arctic Research Laboratory (Barrow), Office of
Naval Research, for their logistical support of fieldwork in summers
1968-1971. Large collections of corals, Foraminifera, and thin sections used in this study were collected
gists in 1959-1964; the collections I
Company, were given
1962 and 1964, while
to the U.S. Geological
given to R. E.
Burton, vice presidents of Shell Oil Company.
Bowsher and Dutro (1957) established the Kayak Shale and
Lisburne Group for exposures near Shainin Lake, Endicott
and published the first major study of the Carboniferous
stratigraphy and faunas. They gave (p. 3) an excellent account of
the earlier history of studies and paleontology of the Carboniferous
rocks of arctic Alaska. Helen
detailed study of Lisburne corals from the Endicott Mountains.
She illustrated and
species of solitary rugose
tabulate corals of Early and Late Mississippian age. She also recognized and illustrated most of the colonial lithostrotionoids that oc-
cur in the Lisburne Group.
Corals are abundant in
of the Carboniferous Lisburne
Brooks Range. These exposures (Text-figs.
2) extend from the Canadian border on the east to the Chukchi Sea
on the west. The study of the Lisburne coral faunas has been made
conjunction with detailed analyses of Lisburne Group biostrati-
graphy, petrology, diagenesis, and environments of deposition.